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Parallel storylines follow English perfume saleswoman Alice Chenery and American comedy writer Gil Raymond, who are seemingly perfect soulmates but never meet, as they deal with their chaotic lives and relationships.
Alice worries that her manager may be after more than a business meeting when she's summoned to the woman's hotel room, Cleo obsesses over a bad review and Milly is strangely drawn to a man's shadow.
Alice bumps into what may be the "Perfect Man" at a cash machine, but their first date includes meeting the family and a spectre from his past. Elsewhere, Cleo and Milly devise a test for Darren's love of Milly.
Alice's yearly lunch with a college chum is interrupted by an obsessed student; Milly and Cleo try out the speed dating fad.
Alice's encounter with the anti-terrorism unit of the local police may have something to do with the romantic deceit she engaged in with her latest beau, and Cleo indulges in some sexual games of her own.
Alice is swept into the posh life of Matthew Chater; Milly and Cleo find themselves arguing over a tee-shirt, a man, and a funeral.
Cleo and Milly get in trouble when they access personal health records at the store's chemist in an effort to screen potential dates for Alice; meanwhile, Alice's boyfriend's job may be its own deterrent.
Alice's finances force her to finally let her Brighton flat and look for something cheaper in London.
Everything seems to be going well with Alice's new bed-in, Douglas, but a message from his ex-girlfriend will plunge them into another round of humiliations and disappointment.
Alice is excited by her opportunity to see a play directed by a comedy actor she admires.
Alice's life is in turmoil, although her housing situation has stabilised, and she wonders if her choice about Douglas was correct.
Getting used to a new flatmate is difficult, unfortunately Alice's date provides no distraction as it descends into the drama that's typical of her outings.
Alice's loneliness becomes pervasive. Is the solution another date, or will the grand plan finally be revealed?
